For anaphase to begin, which of the following must occur?

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	Chromatids must lose their kinetochores.
	Cohesin must attach the sister chromatids to each other.
	Cohesin must be cleaved enzymatically.
	Kinetochores must attach to the metaphase plate.
	Spindle microtubules must begin to depolymerize.

For anaphase to begin, which of the following must occur?

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